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Important Update:

The University of Central Oklahoma physical campus will be closed to all but essential employees through May 31.
Alternative instructional delivery methods will be used for the rest of the spring semester, May intersession courses and summer blocks I and II. Many university services will continue to be available online. For more information, visit the university's COVID-19 website.

Visiting Assistant Professor (Temp)

University of Central Oklahoma

Education and Certifications

B.A. Oberlin College, Oberlin OH

Ph.D. University of Georgia, Athens GA in Botany


University of Central Oklahoma (UCO), Biology, 2015-current

Oklahoma State University (OSU), Botany, 2014-2015

National Science Foundation, Division of Environmental Biology, Evolutionary Processes and Systematics Clusters 2010-2014

Rancho Santa Ana Botanic Garden and Claremont Graduate University (RSA), Program in Botany, 1995-2010

Classes Taught

At UCO: General Biology, Biology: Principles, Biology: Diversity, Cell and Molecular Biology, Genetics

At OSU: Introductory Biology, Introductory Botany, Plant Physiology, Genetics

At RSA: Plant Molecular Biology, Advanced Population Genetics, Molecular Evolution


Plant Adaptation to Novel Environments


Population Genetics

Island Biogeography

Research, Published Work, and Scholarly Activities

Friar EA, McGlaughlin ME, Huang DI, Baldwin BG. In prep. Species tree estimation in a rapid adaptive radiation; an example from the Hawaiian silversword alliance. To be submitted to Evolution.

Friar EA, McGlaughlin ME, Cruse-Sanders JM, Huang DI. Submitted. Fine-scale genetic structure in a hybrid zone between ecologically-divergent members of the Hawaiian silversword alliance, Dubautia arborea and D. ciliolata. Submitted to International Journal of Plant Sciences.

Friar EA. 2017. Review of Plant Cells and Their Organelles. Crop Science 57: 1-2.

Garcia-Verdugo C, Friar EA, Santiago LS. 2013. Ecological role of hybridization in adaptive radiations: a case study in the Dubautia arborea-D. ciliolata (Asteraceae) complex. International Journal of Plant Sciences 174: 749-759.

Cruse-Sanders JM, Parker KC, Friar EA, Huang DI, Mashayekhi S, Prince LM, Otero-Arnaiz A, Casas A. 2013. Managing diversity: Domestication and gene flow in Stenocereus stellatus Riccob. (Cactaceae) in Mexico. Ecology and Evolution 3: 1340-1355.

Huang DI, Friar EA. 2011. Relationships in the Lupinus albifrons species complex (Fabaceae) based on two highly variable chloroplast regions. Systematic Botany 36: 362-370.

McGlaughlin ME, Friar EA. 2011. Evolutionary diversification and geographical isolation in Dubautia laxa (Asteraceae), a widespread member of the Hawaiian silversword alliance. Annals of Botany 107: 357-370.

Baldwin BG, Friar EA. 2010. Dubautia carrii and D. hanaulaensis, new species of the Hawaiian silversword alliance (Compositae, Madiinae) from Molokai and Maui. Novon 20: 1-8.

Friar EA, Prince LM, Cruse-Sanders JM, McGlaughlin ME, Butterworth C, Baldwin, BG. 2007. Hybrid origin and genomic mosaicism of Dubautia scabra (Hawaiian Silversword Alliance; Asteraceae--Madiinae). Systematic Botany, 33: 589-597.

Friar EA, Cruse-Sanders J, McGlaughlin ME. 2007. Gene flow in Dubautia arborea and D. ciliolata: the roles of ecology and isolation by distance in maintaining species boundaries. Molecular Ecology 16: 4028-4038.

Kinney, MS, Columbus, JT, Friar EA. 2007. Unisexual flower, spikelet, and inflorescence development in monoecious/dioecious Bouteloua dimorpha (Poaceae, Chloridoideae). American Journal of Botany 95:123-132.

Lawton-Rauh A, Friar EA, Remington DL. 2007. Collective evolution processes and the tempo of lineage divergence in the Hawaiian silversword alliance adaptive radiation (Heliantheae, Asteraceae). Molecular Ecology 16: 3993-3994.

Friar EA, Prince LM, Roalson EH, McGlaughlin ME, Cruse-Sanders J, DeGroot SJ, Porter JM. 2006. Ecological speciation in the East Maui-endemic Dubautia (Asteraceae) species. Evolution 60: 1777-1792.

McGlaughlin ME, Friar EA. 2007. Clonality in the endangered Ambrosia pumila (Asteraceae) inferred from RAPD markers; implications for conservation and management. Conservation Genetics 8: 319-330.

LaDoux T, Friar EA. 2006. Late-acting self-incompatibility in Ipomopsis tenuifolia (Gray) V. Grant (Polemoniaceae). International Journal of Plant Sciences 167: 463-471

Roalson EH, Friar EA. 2004. Phylogenetic analysis of the nuclear alcohol dehydrogenase (Adh) gene family in Carex section Acrocystis (Cyperaceae) and combined analyses of Adh and nuclear ribosomal ITS and ETS sequences for inferring species relationships. Mol. Phyl. Evol. 33: 671-686.

Kinney MS, Columbus JT, Friar EA. 2003. Molecular evolution of the sex-determining gene TASSELSEED2 in Bouteloua (Poaceae). Mol. Phyl. Evol., 29:519-528.

Roalson EH, Friar EA. 2003. Phylogenetic relationships and biogeographic patterns in North American members of Carex section Acrocystis (Cyperaceae) using nrITS and ETS sequence data. Plant Syst. Evol. 243:175-187.

Friar EA, LaDoux T. 2002. Genetic control of self-incompatibility in Centromadia (Hemizonia) pungens subsp. laevis (Madiinae: Asteraceae). Aliso, 21:1-6.

Friar EA, Boose DL, LaDoux T, Roalson EH, Robichaux RH. 2001. Population structure in the endangered Mauna Loa silversword, Argyroxiphium kauense (Asteraceae), and its bearing on reintroduction. Molecular Ecology, 10: 1657-1664.

Roalson EH, Columbus JT, Friar EA. 2001. Phylogenetic relationships in Cariceae (Cyperaceae) based on ITS (nrDNA) and trnT-L-F (cpDNA) region sequences: assessment of subgeneric and sectional relationships in Carex with emphasis on section Acrocystis. Syst. Bot. 26: 318-341.

Barrier M, Friar EA, Robichaux R, Purugganan M. 2000. Interspecific evolution in plant microsatellite structure. Gene 241: 101-105.

Friar EA, LaDoux T, Roalson EH, Robichaux RH. 2000. Microsatellite analysis of a population crash and bottleneck in the Mauna Kea silversword (Argyroxiphium sandwicense ssp. sandwicense), and its implications for reintroduction. Molecular Ecology 9: 2027-2034.

Morrell PL, Porter JM, Friar EA. 2000. Intercontinental dispersal: the origin of the widespread South American plant species Gilia laciniata (Polemoniaceae) from a rare California and Oregon coastal endemic. Plant Syst. Evol. 224: 13-32.

Roalson EH, Friar EA. 2000. Supraspecific classification of Eleocharis (Cyperaceae)

Books and Book Chapters

Roalson EH, Friar EA. 2000. Supraspecific classification of Eleocharis (Cyperaceae) revisited: evidence from the Internal Transcribed Spacer (ITS) region of nuclear ribosomal DNA. Systematic Botany 25: 323-336.

Ashworth VATM, O'Brien BC, Friar EA. 1999. Fingerprinting Juniperus communis L. cultivars using RAPD markers. Madroño 46: 134-141.

Robichaux RH, Friar EA, Mount DW. 1997. Molecular genetic consequences of a population bottleneck associated with reintroduction of the Mauna Kea silversword (Argyroxiphium sandwicense ssp. sandwicense [Asteraceae]). Conservation Biology 11: 1140-1146.

Friar EA, Robichaux RH, Mount DW. 1996. Molecular genetic variation following a population crash in the endangered Mauna Kea silversword, Argyroxiphium sandwicense ssp. sandwicense (Asteraceae). Molecular Ecology 5: 687-691.

Friar EA, Falk M, Mount DW. 1995. Use of random amplified polymorphic DNA (RAPD) markers for genetic analysis of Lilium parryi, a rare Arizona plant. In: Maschinski J, Hammond HD, Holter L (eds.) Southwestern Rare and Endangered Plants: Proceedings of the Second Conference.

Friar EA, Kochert G. 1994. A study of genetic variation and phylogeny of the genus Phyllostachys (Bambusoideae: Poaceae) using nuclear restriction fragment length polymorphisms. Theoretical and Applied Genetics 89: 265-270

Friar EA, Kochert G. 1991. Bamboo germplasm screening with nuclear restriction fragment length polymorphisms. Theoretical and Applied Genetics 82: 697-703.

Columbus JT, Friar EA, Porter JM, Prince LM, Simpson MG (eds.). 2007. Monocots: Comparative biology and evolution. 2 vols. Rancho Santa Ana Botanic Garden, Claremont, California, USA.

Kinney, MS, Columbus JT, Friar EA. 2007. Evolution of development of unisexual and hermaphrodite flowers in Bouteloua (Poaceae). Pp. 605-614 In: Columbus JT, Friar EA, Porter JM, Prince LM, Simpson MG (eds.), Monocots: Comparative biology and evolution. 2 vols. Rancho Santa Ana Botanic Garden, Claremont, California, USA.

Friar EA. 2005. Isolation of DNA from plants with large amounts of secondary metabolites. In: Zimmer EA, Roalson EH (eds.) Molecular Evolution: Producing the Biochemical Data, Part B. Elsevier Press, pp. 3-8.

Friar EA, Robichaux RH. 2003. Conservation genetics of the silversword alliance. In: Carlquist S, Baldwin BG, Carr GD (eds.) Tarweeds and Silverswords: Evolution of the Madiinae. Missouri Botanical Garden Press, St. Louis, MO., pp.183-192.

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